© R. P. Anderson 

Copyright © 2004-2017.

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Peterson, A.T., J. Soberón, R. G. Pearson, R. P. Anderson, E. Martínez-Meyer, M. Nakamura, and M. B. Araújo. 2011. Ecological niches and geographic distributions. Monographs in Population Biology, 49. Princeton University Press.





Book chapters:


Anderson, R. P. 2015. Family Heteromyidae. Pp. 51-58 in: Mammals of South America, Volume 2: Rodents. J. L. Patton, U. F. J. Pardiñas, and G. D’Elía (editors). University of Chicago Press. pdf




Journal publications:


Araújo, M. B., R. P. Anderson, A. M. Barbosa, C. M. Beale, C. F. Dormann, R. Early, R. A. Garcia, A. Guisan, L. Maiorano, B. Naimi, R. B. O’Hara, N. E. Zimmermann, and C. Rahbek. 2019. Standards for models in biodiversity assessments. Science Advances, 5: eaat4858. pdf (Online Supplementary Material)

Bohl, C. L., J. M. Kass, and R. P. Anderson. 2019. A new null model approach to quantifying performance and significance for ecological niche models of species distributions. Journal of Biogeography. 46:1101–1111. [Editor’s Choice]. pdf (Online Supplementary Material)


Peterson, A. T. et al. (55 total authors). 2019. Open access solutions for biodiversity journals: Do not replace one problem with another. Diversity and Distributions, 25:5–8. pdf

Peterson, A. T., R. P. Anderson, M. E. Cobos, M. Cuahutle, A. P. Cuervo-Robayo, L. E. Escobar, M. Fernandez, D. Jiménez-García, A. Lira-Noriega, J. M. Lobo, F. Machado-Stredel, E. Martínez-Meyer, C. Nuñez-Penichet, J. Nori, L. Osorio-Olvera, M. T. Rodríguez, O. Rojas-Soto, D. Romero-Álvarez, J. Soberón, S. Varela, and C. Yañez-Arenas. 2019. Curso Modelado de Nicho Ecológico, Versión 1.0. Biodiversity Informatics, 14:1–7. pdf


Galante, P. J., B. Alade, R. Muscarella, S. A. Jansa, S. M. Goodman, and R. P. Anderson. 2018. The challenge of modeling niches and distributions for data-poor species: a comprehensive approach to model complexity. Ecography. 41:726–736. pdf

Gerstner, B. E., J. M. Kass, R. Kays, K. M. Helgen, and R. P. Anderson. 2018. Revised distributional estimates for the recently discovered olinguito, (Bassaricyon neblina), with comments on natural and taxonomic history. Journal of Mammalogy. 99:321-332. pdf

Guevara, L., B. E. Gerstner, J. M. Kass, and R. P. Anderson. 2018. Toward ecologically realistic predictions of species distributions: A cross-time example from tropical montane cloud forests. Global Change Biology. 24:1511-1522. pdf

Guevara, L., L. León-Paniagua, J. Rios, and R. P. Anderson. 2018. Variation among Global Circulation Models for reconstructions of geographic distributions at the Last Glacial Maximum: relevance for phylogeography. Ecosistemas. 27:62-76. pdf

Kass, J. M., B. Vilela, M. E. Aiello-Lammens, R. Muscarella, C. Merow, and R. P. Anderson. 2018. WALLACE: A flexible platform for reproducible modeling of species niches and distributions built for community expansion. Methods in Ecology and Evolution.

9:1151-1156. pdf

Sanín, C. and R. P. Anderson. 2018. A framework for simultaneous tests of abiotic, biotic, and historical drivers of species distributions: empirical tests for North American wood-warblers based on climate and pollen. American Naturalist. 192:E48-E61. pdf


Anderson, R. P. 2017. When and how should biotic interactions be considered in models of species niches and distributions? Journal of Biogeography. 44:8-17.​ pdf

Boria, R. A., L. E. Olson, S. M. Goodman, and R. P. Anderson. 2017. A single-algorithm ensemble approach to estimating suitability and uncertainty: cross-time projections for four Malagasy tenrecs. Diversity and Distributions. 23:196-208. pdf (Online Supplementary Material)

Phillips, S. J., R. P. Anderson, M. Dudík, R. E. Schapire, M. E. Blair. 2017. Opening the black box: an open–source release of Maxent. Ecography. 40:887-893. pdf



Cook J. A., S. E. Greiman, S. J. Agosta, R. P. Anderson, B. S. Arbogast, R. J. Baker, W. Boeger, R. D. Bradley, D. R. Brooks, R. Cole, J. R. Demboski, A. P. Dobson, J. L. Dunnum, R. P. Eckerlin, J. Esselstyn, K. E. Galbreath, J. Hawdon, H. E. Hoekstra, S. J. Kutz, J. E. Light, L. E. Olson, B. D. Patterson, J. L. Patton, A. J. Phillips, E. Rickart, D. S. Rogers, M. E. Siddall, V. V. Tkach, E. P. Hoberg. 2016. Transformational principles for NEON sampling of mammalian parasites and pathogens: a response to Springer and colleagues. BioScience. 66:917-919. pdf


Soley-Guardia, M., E. E. Gutiérrez, D. M. Thomas, J. Ochoa-G., M. Aguilera, and R. P. Anderson. 2016. Are we overestimating the niche? Removing marginal localities helps ecological niche models detect environmental barriers. Ecology and Evolution. 6:1267-1279. pdf (Online Supplementary Material)


Aiello-Lammens, M. E., R. A. Boria, A. Radosavljevic, B. Vilela, and R. P. Anderson. 2015. spThin: an R package for spatial thinning of species occurrence records for use in ecological niche models. Ecography. 38:541-545. pdf

Anderson, R. P. 2015. El modelado de nichos y distribuciones: no es simplemente “clic, clic, clic.” [With English and French translations: Modeling niches and distributions: it’s not just “click, click, click” and La modélisation de niche et de distributions: ce n’est pas juste “clic, clic, clic”]. Biogeografía. 8:4-27. [invited by the editors; not peer-reviewed] pdf


Anderson, R. P. 2014. [Review of:] Ponsà Fontanals, M. 2011. Osvaldo Reig: la vida itinerante de un biólogo evolucionista. Journal of Mammalogy. 95:187–188. pdf [not peer-reviewed]


Boria, R. A., L. E. Olson, S. M. Goodman, and R. P. Anderson. 2014. Spatial filtering to reduce sampling bias can improve the performance of ecological niche models. Ecological Modelling. 275:73-77. pdf


Gutiérrez, E. E., R. P. Anderson, R. S. Voss, J. Ochoa-G., M. Aguilera, and S. A. Jansa. 2014. Phylogeography of the mouse opossum Marmosa robinsoni: with insights into the biogeographic history of dry habitats in northern South America. Journal of Mammalogy. 95:1175-1188. pdf


Gutiérrez, E. E., R. A. Boria, and R. P. Anderson. 2014. Can biotic interactions cause allopatry? Niche models, competition, and distributions of South American mouse opossums. Ecography. 37:741-753. pdf (Online Supplementary Material)


Muscarella, R., P. J. Galante, M. Soley-Guardia, R. A. Boria, J. M. Kass, M. Uriarte, and R. P. Anderson. 2014. ENMeval: An R package for conducting spatially independent evaluations and estimating optimal model complexity for MAXENT ecological niche models. Methods in Ecology and Evolution. 5:1198-1205. pdf


Radosavljevic, A. and R. P. Anderson. 2014. Making better Maxent models of species distributions: complexity, overfitting, and evaluation. Journal of Biogeography. 41:629-643. pdf (Online Supplementary Material)


Soley-Guardia, M., A. Radosavljevic, J. L. Rivera, and R. P. Anderson. 2014. The effect of spatially marginal localities in modelling species niches and distributions. Journal of Biogeography. 41:1390-1401. pdf (Online Supplementary Material S1, Online Supplementary Material S2)


Varela, S., R. P. Anderson, R. García-Valdés, and F. Fernández-González. 2014. Environmental filters reduce the effects of sampling bias and improve predictions of ecological niche models. Ecography. 37:1084–1091. pdf


Waltari, E., Schroeder, R., Mcdonald, K., Anderson, R.P., & Carnaval, A. 2014. Bioclimatic variables derived from remote sensing: Assessment and application for species distribution modelling. Methods in Ecology and Evolution. 1033-1042. pdf



Anderson, R. P. 2013. A framework for using niche models to estimate impacts of climate change on species distributions. Annals of the New York Academy of Sciences. 1297:8-28. pdf


Shcheglovitova, M. and R. P. Anderson. 2013. Estimating optimal complexity for ecological niche models: a jackknife approach for species with small sample sizes. Ecological Modelling. 269:9-17. pdf (Online Supplementary Material)



Anderson, R. P., E. E. Gutiérrez, J. Ochoa-G., F. J. García, and M. Aguilera. 2012. Faunal nestedness and species-area relationship for small non-volant mammals in “sky islands” of northern Venezuela. Studies on Neotropical Fauna and Environment. 47:157-170. pdf (Online Supplementary Material)


Anderson, R. P. 2012. Harnessing the world’s biodiversity data: promise and peril in ecological niche modeling of species distributions. Annals of the New York Academy of Sciences, 1260:66-80. pdf



Anderson, R. P. and I. Gonzalez, Jr. 2011. Species-specific tuning increases robustness to sampling bias in models of species distributions: an implementation with Maxent. Ecological Modelling, 222:2796-2811. pdf



Anderson, R. P. and A. Raza. 2010. The effect of the extent of the study region on GIS models of species geographic distributions and estimates of niche evolution: preliminary tests with montane rodents (genus Nephelomys) in Venezuela. Journal of Biogeography, 37:1378-1393. pdf



Anderson, R. P. and E. E. Gutiérrez. 2009. Taxonomy, distribution, and natural history of the genus Heteromys (Rodentia: Heteromyidae) in central and eastern Venezuela, with the description of a new species from the Cordillera de la Costa. In: R. S. Voss and M. D. Carleton (editors), Systematic mammalogy: contributions in honor of Guy G. Musser. Bulletin of the American Museum of Natural History, 331:33-93. pdf



Anderson, R. P. and S.A. Jansa. 2007. Genetic comparisons between Heteromys desmarestianus and the recently described H. nubicolens (Rodentia: Heteromyidae) in northwestern Costa Rica. Mammalian Biology, 72:54-61. pdf



Anderson, R. P., A. T. Peterson, and S. L. Egbert. 2006. Vegetation-index models predict areas vulnerable to purple loosestrife (Lythrum salicaria) invasion in Kansas. Southwestern Naturalist, 51:471-480. pdf


Anderson, R. P. and R. M. Timm. 2006. A new montane species of spiny pocket mouse (Rodentia: Heteromyidae: Heteromys) from northwestern Costa Rica. American Museum Novitates, 3509:1-38. pdf


Anderson, R. P., M. Weksler, and D. S. Rogers. 2006. Phylogenetic analyses of spiny pocket mice (Heteromyidae: Heteromyinae) based on allozymic and morphological data. Journal of Mammalogy, 87:1218-1233. pdf


Elith, J., C. H. Graham, R. P. Anderson, M. Dudík, S. Ferrier, A. Guisan, R. J. Hijmans, F. Huettmann, J. R. Leathwick, A. Lehmann, J. Li, L. G. Lohmann, B. A. Loiselle, G. Manion, C. Moritz, M. Nakamura, Y. Nakazawa, J. M. Overton, A. T. Peterson, S. J. Phillips, K. Richardson, R. Scachetti-Pereira, R. E. Schapire, J. Soberón, S. Williams, M. S. Wisz, and N. E. Zimmerman. 2006. Novel methods improve prediction of species’ distributions from occurrence data. Ecography, 29:129-151. pdf

[This work was featured in the "Research Highlights" section of Nature, 18 May 2006: 259 pdf].


Phillips, S. J., R. P. Anderson, and R. E. Schapire. 2006. Maximum entropy modeling of species geographic distributions. Ecological Modelling, 190:231-259. pdf



Anderson, R. P. and Martínez-Meyer. 2004. Modeling species’ geographic distributions for preliminary conservation assessments: an implementation with the spiny pocket mice (Heteromys) of Ecuador. Biological Conservation, 116:167-179. pdf


Gómez-Laverde, M., R. P. Anderson, and L. F. García. 2004. Integrated systematic reevaluation of the Amazonian genus Scolomys (Rodentia: Sigmodontinae). Mammalian Biology, 69:119-139. pdf



Anderson, R. P. 2003. Real vs. artefactual absences in species distributions: tests for Oryzomys albigularis (Rodentia: Muridae) in Venezuela. Journal of Biogeography, 30:591-605. pdf


Anderson, R. P. 2003. Taxonomy, distribution, and natural history of the genus Heteromys (Rodentia: Heteromyidae) in western Venezuela, with the description of a dwarf species from the Península de Paraguaná. American Museum Novitates, 3396:1-43. pdf


Anderson, R. P., D. Lew, and A. T. Peterson. 2003. Evaluating predictive models of species' distributions: criteria for selecting optimal models. Ecological Modelling, 162:211-232. pdf



Anderson, R. P., M. Gómez-Laverde, and A. T. Peterson. 2002. Geographical distributions of spiny pocket mice in South America: insights from predictive models. Global Ecology & Biogeography, 11:131-141. pdf


Anderson, R. P. and C. O. Handley, Jr. 2002. Dwarfism in insular sloths: biogeography, selection, and evolutionary rate. Evolution, 56:1045-1058. pdf


Anderson, R. P. and P. Jarrín-V. 2002. A new species of spiny pocket mouse (Heteromyidae: Heteromys) endemic to western Ecuador. American Museum Novitates, 3382:1-26. pdf


Anderson, R. P., A. T. Peterson, and M. Gómez-Laverde. 2002. Using niche-based GIS modeling to test geographic predictions of competitive exclusion and competitive release in South American pocket mice. Oikos, 98:3-16. pdf



Anderson, R. P. and C. O. Handley, Jr. 2001. A new species of three-toed sloth (Mammalia: Xenarthra) from Panamá, with a review of the genus Bradypus. Proceedings of the Biological Society of Washington, 114:1-33. pdf


[This work was featured in the "Discoveries" section of BBC Wildlife magazine, October 2001:17; and in the “Wildlife” section of National Geographic magazine, March 2006:2, 20].

Download Excel file of Bradypus localities from this paper



Anderson, R. P. 1999 [2000]. Preliminary review of the systematics and biogeography of the spiny pocket mice (Heteromys) of Colombia. Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales, 23 (Suplemento especial):613-630. pdf



Anderson, R. P. and P. J. Soriano. 1999. The occurrence and biogeographic significance of the southern spiny pocket mouse Heteromys australis in Venezuela. Zeitschrift für Säugetierkunde, 64:121-125. pdf


Lim, B. K., M. D. Engstrom, R. M. Timm, R. P. Anderson, and L. C. Watson. 1999. First records of 10 bat species in Guyana and comments on diversity of bats in Iwokrama Forest. Acta Chiropterologica, 1:179-190. pdf



Cadena, A., R. P. Anderson, and P. Rivas-Pava. 1998. Colombian mammals from the Chocoan slopes of Nariño. Occasional Papers of the Museum, Texas Tech University, 180:1-15. pdf



Anderson, R. P. 1994. Faunal Analysis of Feature 4, 14JW1. In: B. Logan and W. E. Banks (editors), White Rock revised: archaeological investigation of the White Rock and Warne sites, Lovewell Reservoir, Jewell County, Kansas. University of Kansas Museum of Anthropology Project Report Series, 85: 55-64. [invited by the editors; not peer-reviewed]






















Ph.D. dissertation:


Anderson, R. P. 2001. Systematics and biogeographic modeling of spiny pocket mice (Rodentia: Heteromyidae: Heteromys) in South America. Ph.D. Dissertation, University of Kansas, Lawrence. ix + 169 pp.




Publications (R. P. Anderson)

Quindío, Colombia, 1995

Western Nariño, Colombia in 1995; see Cadena et al. 1998